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These phosphorylated R-SMADs dissociate from the receptor and form a trimeric complex with SMAD4, which translocates into the nucleus to regulate transcription of fibrosis-related genes, including -smooth muscle actin (-SMA) and collagen I [40, 41]

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To summarize, there is evidence to support LC-FFA induced ER stress in human -cells although this pathway has mainly been characterized in rodents